Mission 13: Genesis
Tilt Skull – Once again you’ll have to search the area where the Warden appears. In this case, it will be the location where the very first one appears. There will be some rocks nearby for you to search in order to locate this Skull.
Mission 14: The Breaking
Famine Skull – Wardens appear to be a trend when it comes to locating skulls. In the middle of The Breaking a Warden will appear on a platform. Check underneath that platform to locate this Skull (you may want to clear out the enemies first).
Mission 15: Guardians
Tough Luck Skull – While progressing through the mission as normal you’ll eventually reach a ledge that looks over the Gamma Core. Look around this area to locate your final Skull in Halo 5: Guardians. Jump to the other side of the Pelican wreckage near the cliff to locate this beauty.